Boundaries: Punctuation and Sentence Structure

SAT Reading and Writing· difficulty 4/5

The botanist's discovery transformed our understanding of ancient plant biology ______ fossilized leaves contained intact chemical signatures of photosynthesis from 200 million years ago.

Which choice completes the text so that it conforms to the conventions of Standard English?

  • A

    ;

  • B

    :

    check_circle
  • C

    (empty)

  • D

    , which

Explanation

A colon introduces an explanation of how the discovery transformed understanding. A semicolon could work but a colon better signals explanation; " " is a run-on; ", which" creates an unclear modifier.

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